» Dr. Douglas Fudge
Associate Professor

Schmid College of Science and Technology; Biological Sciences
Dr. Douglas Fudge
Office Location:
Hashinger Science Center 308
Cornell University, Bachelor of Arts
Cornell University, Master of Arts
University of Guelph, Master of Science
The University of British Columbia, Ph.D.

Research Interests

Dr. Fudge’s research aims to understand the biophysics of marine animals, with a focus on processes such as predator defense, feeding, and locomotion. Current projects aim to understand predator-prey interactions between hagfishes and sharks, the function and biogenesis of hagfish slime, and the development of biomimetic applications inspired by hagfish slime.


marine biology, biomaterials, biomechanics, animal physiology, hagfish, slime, keratins, animal defense

Recent Creative, Scholarly Work and Publications
Turko AJ, Kültz D, Fudge DS, Croll RP, Smith FM, Stoyek MR, and Wright PA (2017). Skeletal stiffening in an amphibious fish out of water is a response to increased body weight. Journal of Experimental Biology, accepted 08/07/17.
Freedman C and Fudge DS (2017). Hagfish Houdinis: biomechanics and behavior of squeezing through small openings. Journal of Experimental Biology 220: 822-7.
Soomro A, Alsop RJ, Negishi A, Kreplak L, Fudge DS, Kuczmarski ER , Goldman RD, Rheinstadter MC (2017) Giant axonal neuropathy alters the structure of keratin intermediate filaments in human hair. J Roy Soc Interface 14 20170123.
Fudge DS, Winegard TM, Herr JE (2016). Hagfish slime and slime glands. In Phylogeny, Anatomy and Physiology of Ancient Fishes, Zaccone G, Dabrowski K, de Oliveira Fernandes JM, and Hedrick M (eds) 272-90.
Fudge DS and Schorno S (2016). The hagfish gland thread cell: a fiber-producing cell involved in predator defense. Cells. 5: 25.
Icardo JM, Colvee E, Schorno S, Lauriano ER, Fudge DS, Glover CN, and Zaccone G (2016). Morphological analysis of the hagfish heart. I. The ventricle, the arterial connection and the ventral aorta. Journal of Morphology, 277: 326-40.
Icardo JM, Colvee E, Schorno S, Laureano ER, Fudge DS, Glover CN, and Zaccone G (2016). Morphological analysis of the hagfish heart. II. The venous pole and the pericardium. Journal of Morphology, 277: 853–865.
Weatherbee-Martin N, Xu L, Hupe A, Kreplak L, Fudge DS, Liu XQ and Rainey JK (2016). Identification of wet-spinning and post-spin stretching methods amenable to recombinant spider aciniform silk. Biomacromolecules, 17: 2737-46.
Fudge DF, Herr JE, and Winegard TM (2015) Hagfish slime: origins, functions, and mechanisms. In Hagfish Biology. Goss GG and Edwards S (eds), 321.
Zaccone G, Fudge D, Winegard T, Capillo G, Kuciel M, Funakoshi K and Lauriano E (2015). Confocal imaging and phylogenetic considerations of the subcutaneous neurons in the Atlantic hagfish Myxine glutinosa. Acta Zoologica. 96: 209-217.
Fudge DS, Schorno S, Ferraro S (2015). Physiology, biomechanics, and biomimetics of hagfish slime. Annual Review of Biochemistry. 84: 947-67.
Won G-J, Fudge DS, Choh V (2015). The effects of actomyosin disruptors on the mechanical integrity of the avian crystalline lens. Molecular Vision. 21: 98-109.
Fudge DS (2014) Fifty years of JR Platt’s “Strong Inference”. Journal of Experimental Biology 217: 1202-4.
Winegard TM, Herr JE, Mena C, Lee B, Dinov I, Bird D, Bernards M, Hobel S, Van Valkenburgh B, Toga A & Fudge DS (2014). Coiling and maturation of a high performance fibre in hagfish slime gland thread cells, Nature Communications 5: 3534.
Herr JE, Clifford A, Goss GG, and Fudge DS (2014). Defensive slime formation in Pacific hagfish requires Ca2+ and aquaporin mediated swelling of released vesicles, Journal of Experimental Biology 217: 2288-96.
Bernards MA, Oke I, Heyland A and Fudge DS (2014). Spontaneous unraveling of hagfish slime thread skeins is mediated by a seawater- soluble protein adhesive, Journal of Experimental Biology 217: 1263-68.
Pinto N, Negishi A, Yang F-C, Rheinstadter M, Gillis TE, and Fudge DS (2014). Self-assembly enhances the strength of fibres made from vimentin intermediate filament proteins, Biomacromolecules 15: 574-581.
Gleason JE, Fudge DS and Robinson B (2014). Eco-mechanics of lamellar autotomy in larval damselflies, Journal of Experimental Biology 217: 185-91.
Shadwick RE, Schiller LL, and Fudge DS (2013). Physiology of swimming and migration in tunas. In Swimming Physiology of Fish, Palstra AP and Planas JV (eds.), Springer-Verlag, pp 45-78.