Dr. Douglas Fudge

Dr. Douglas Fudge

Schmid College of Science and Technology; Biological Sciences
Expertise: Marine Biology; Biomaterials; Biomechanics; Animal Physiology; Hagfish; Slime;
Office Location: Keck Center for Science and Engineering 265
Phone: (714) 628-7213
Scholarly Works:
Digital Commons
Cornell University, Bachelor of Arts
Cornell University, Master of Arts
University of Guelph, Master of Science
The University of British Columbia, Ph.D.


Research Interests

Dr. Fudge’s research aims to understand the biophysics of marine animals, with a focus on processes such as predator defense, feeding, and locomotion. Current projects aim to understand predator-prey interactions between hagfishes and sharks, the function and biogenesis of hagfish slime, and the development of biomimetic applications inspired by hagfish slime.


marine biology, biomaterials, biomechanics, animal physiology, hagfish, slime, keratins, animal defense

Recent Creative, Scholarly Work and Publications

Zeng Y, Petrichko S, Nieders K, Plachetzki D, Fudge D. Evolution of a remarkable intracellular polymer and extreme cell allometry in hagfishes. Current Biology. 2021 Nov 22;31(22):5062-8.
Bressman N and Fudge DS (2021). From reductionism to synthesis: The case of hagfish slime. Comparative Biochemistry and Physiology Part B: Biochemistry and Molecular Biology, 110610.
Mincarone MM, Plachetzki D, McCord CL, Winegard TM, Fernholm B, Gonzalez CJ, Fudge DS (2021) Review of the hagfishes (Myxinidae) from the Galapagos Islands, with descriptions of four new species and their phylogenetic relationships. Zoological Journal of the Linnean Society, 192: 453-474.
Fudge DS and Turko AJ (2020) The best predictions in experimental biology are critical and persuasive. Journal of Experimental Biology, 223: jeb231894.
Cerullo A, Allam B, Baer A, Barnes WJP, Barrientos Z, Deheyn DD, Fudge DS, Gould J, Harrington MJ, Holford M, Hung C-S, Jain G, Lai T-Y, Mayer G, Medina M, Monge-Nájera J, Napolitano T, Espinosa EP, Schmidt S, Thompson EM, Braunschweig AB (2020) Comparative animal mucomics: Inspiration for functional materials from a ubiquitous and understudied biopolymer. ACS Biomaterials Science & Engineering, 6: 5377-5398.
Fudge DS, Ferraro SN, Siwiecki SA, Hupé A, and Jain G (2020). A new model of hagfish slime mucous vesicle stabilization and deployment. Langmuir, 36: 6681–6689.
McCord C, Whiteley E, Liang J, Trejo C, Caputo R, Itehua E; Hasan H, Hernandez S, Jagnandan K, Fudge DS (2020). Dose effects of three common fish anesthetics on Pacific hagfish (Eptatretus stoutii). Fish Physiology and Biochemistry, Jan 18: 1-13.
Black KL, Fudge D, Jarvis, WM, & Robinson BW (2019). Functional plasticity in lamellar autotomy by larval damselflies in response to predatory larval dragonfly cues. Evolutionary Ecology, 33, 257-272.
Jain G, Starksen M, Singh K, Hoang C, Yancey P, McCord C, and Fudge DS (2019). High concentrations of trimethylamines in slime glands inhibit skein unraveling in Pacific hagfish. Journal of Experimental Biology 222, doi: 10.1242/jeb.213793.
Schorno S, Gillis TE, Fudge DS (2018) Cellular mechanisms of slime gland refilling in Pacific hagfish (Eptatretus stoutii). Journal of Experimental Biology jeb-183806.
Schorno S, Gillis TE, Fudge DS (2018) Emptying and refilling of slime glands in Atlantic (Myxine glutinosa) and Pacific (Eptatretus stoutii) hagfishes. Journal of Experimental Biology jeb-172254.
Chaudhary G, Fudge DS, Macias- Rodriguez; Ewoldt RH (2018) Concentration-independent mechanics and structure of hagfish slime. Acta Biomaterialia, https://doi.org/10.1016/j.actbio.2018.08.022
Bernards Jr., MA, Schorno S, McKenzie E, Winegard, TM, Oke I, Plachetzki D & Fudge, DS (2018). Unraveling inter-species differences in hagfish slime skein deployment. Journal of Experimental Biology, 221, 1-11.
Turko AJ, Kültz D, Fudge DS, Croll RP, Smith FM, Stoyek MR, and Wright PA (2017). Skeletal stiffening in an amphibious fish out of water is a response to increased body weight. Journal of Experimental Biology 220:3621-31.
Freedman C and Fudge DS (2017). Hagfish Houdinis: biomechanics and behavior of squeezing through small openings. Journal of Experimental Biology 220: 822-7.
Soomro A, Alsop RJ, Negishi A, Kreplak L, Fudge DS, Kuczmarski ER , Goldman RD, Rheinstadter MC (2017) Giant axonal neuropathy alters the structure of keratin intermediate filaments in human hair. J Roy Soc Interface 14 20170123.
Boggett S, Stiles JL, Summers AP, Fudge DS (2017) Flaccid skin protects hagfishes from shark bites. Journal of The Royal Society Interface 14:20170765.